Jean de Villiers will face Samoa in Pretoria.
Springbok coach Heyneke Meyer gave De Villiers until 24 hours before the match to confirm his availability and the player will start at inside centre.
Jan Serfontein trained all week at inside centre and will in all likelihood play the last 30 to 40 minutes.
Morne Steyn again starts ahead of Pat Lambie.
Willem Alberts was also given until the captain’s run to confirm his fitness and he also survived the session.
Adriaan Strauss retains the run on No 2 jersey while Bismarck du Plessis is nursed back after nine months out of the game.
Boks – 15. Willie le Roux, 14. Bryan Habana, 13. JJ Engelbrecht, 12. Jean de Villiers (captain), 11. Bjorn Basson, 10. Morné Steyn, 9. Ruan Pienaar, 8. Pierre Spies, 7. Willem Alberts, 6. Francois Louw, 5. Flip van der Merwe, 4. Eben Etzebeth, 3. Jannie du Plessis, 2. Adriaan Strauss, 1. Tendai Mtawarira
Substitutes – 16. Bismarck du Plessis, 17. Trevor Nyakane, 18. Coenie Oosthuizen, 19. Juandre Kruger, 20. Siya Kolisi, 21. Piet van Zyl, 22. Pat Lambie, 23. Jan Serfontein