Willem Alberts will start at No 8 against Argentina in Mendoza with Jacques Potgieter at blindside flank.
Keegan Daniel was the Bok eighthman at Newlands but was substituted shortly after half-time and has been relegated to the bench. Adriaan Strauss, as expected, replaces the injured Bismarck du Plessis at hooker, while Tiaan Liebenberg has beaten Craig Burden to jersey No 16.
‘It was a difficult decision between Jacques and Keegan, but we have gone with horses for courses this time,’ said Bok coach Heyneke Meyer. ‘We’re expecting it will be very physical early in the game and it might open up later, which perfectly suits the strongpoints of both Jacques and Keegan.
‘The Pumas also mauled quite well in Cape Town and we were not happy with the defence of their mauls, so with the extra weight Jacques brings, we will have to ensure we stop their drives.’
Bryan Habana injured his left ankle against Argentina in Cape Town and passed a fitness test on Wednesday morning.
‘Having Bryan is obviously a big boost as we’ve lost a number of players with lots of experience after JP Pietersen, Pierre Spies and Bismarck du Plessis already had to withdraw from our squad,’ said Meyer.
South Africa – 15 Zane Kirchner, 14 Bryan Habana, 13 Jean de Villiers (c), 12 Frans Steyn, 11 Lwazi Mvovo, 10 Morné Steyn, 9 Francois Hougaard, 8 Willem Alberts, 7 Jacques Potgieter, 6 Marcell Coetzee, 5 Andries Bekker, 4 Eben Etzebeth, 3 Jannie du Plessis, 2 Adriaan Strauss, 1 Beast Mtawarira.
Subs: 16 Tiaan Liebenberg, 17 Pat Cilliers, 18 Flip van der Merwe, 19 Keegan Daniel, 20 Ruan Pienaar, 21 Pat Lambie, 22 JJ Engelbrecht.